The officiating mistake was a doozy — and it cannot be fixed. The play that followed was even more astonishing.

One of the oddest endings imaginable left host Oklahoma State stunned and Central Michigan celebrating.

A misinterpreted rule extended the game when it should have been over and allowed the Chippewas to score a wild winning touchdown on a Hail Mary and lateral — yes, both — that covered half the field for an astonishing 30-27 upset of No. 22 Oklahoma State on Saturday.

Oklahoma State tried to kill the final four seconds of the game by throwing the ball away on fourth down, but the Cowboys were penalized for intentional grounding, which is a loss-of-down penalty. Rules state that the game cannot end on an accepted live-ball penalty, referee Tim O'Dey of the Mid-American Conference, CMU's league, said.

“There's an exception to the rule that says if enforcement of the foul involves a loss of down, then that brings the game to an end,” O'Dey told a pool reporter.

O'Dey said after conferring with NCAA rules committee secretary Rogers Redding after the game, the crew determined the “extension should not have happened.”

But the final result stood. Article 3b of the NCAA rule book states: When the referee declares that the game is ended, the score is final.

“Despite the error, this will not change the outcome of the contest,” Bill Carollo, the NCAA's coordinator of football officials, said in a statement released by the MAC.

Corey Willis scored the winning touchdown after grabbing a lateral from Jesse Kroll at the Oklahoma State 12 and fighting his sway into the end zone.

Oklahoma State (1-1) thought it had won when Mason Rudolph threw the ball away on fourth down as the clock expired, instead of just taking a knee and risking giving the ball back to Central Michigan, but it backfired. After the officials conferred, they assessed an intentional-grounding penalty and gave Central Michigan another play.

Cooper Rush lofted a Hail Mary pass that hit Kroll just inside the 10. As Kroll was being taken down, he pitched it back to Willis at about the 12, who cut across the field and barely managed to score while being dragged down.

Pittsburgh 42, Penn State 39: The Panthers and Nittany Lions spent the runup to the renewal of their series after a 15-year break debating whether they're still rivals. There's no doubt now.

James Conner ran for 117 yards and a touchdown and caught another and cornerback Ryan Lewis intercepted quarterback Trace McSorley in the end zone with just over a minute to play as Pitt held on.

Fullback George Aston ran for a pair of scores for the Panthers (2-0), who nearly let a 21-point lead slip away only to hold on late when Lewis stepped in front of an underthrown pass by McSorley as the majority of the largest crowd to ever watch a sporting event in Pittsburgh — an announced 69,983 — erupted.

No. 2 Clemson 30, Troy 24: The host Tigers (2-0) survived a host of mistakes — including an embarrassing early celebration punt-return gaffe by Ray-Ray McCloud that cost a touchdown on a dynamic 75-yard run right before halftime — to hold off the Trojans (1-1).

Arkansas 41, No. 15 TCU 38, 2OT: Austin Allen ran 5 yards for the winning score in the second overtime, and the Razorbacks (2-0) pulled out a victory over the host Horned Frogs (1-1).

No. 18 Notre Dame 39, Nevada 10: DeShone Kizer threw two touchdown passes and ran for another score to help the host Fighting Irish (1-1) beat the Wolf Pack (1-1).

Record crowd: The “Battle at Bristol” between No. 17 Tennessee and Virginia Tech at Bristol (Tenn.) Motor Speedway drew an NCAA single-game record crowd of 156,990.