LOS ANGELES — Penn State traveled about 2,500 miles across the continent on its first road trip in the new Big Ten. The Nittany Lions then spent most of Saturday playing from behind against Southern California.

Thanks to several clutch throws by Drew Allar, a few key defensive stops and one of the greatest games by a tight end in FBS history, the Nittany Lions can make the long trip home with their unbeaten record and national championship dreams intact.

Tyler Warren caught 17 passes to tie the FBS single-game record for tight ends, Ryan Barker hit a 36-yard field goal in overtime and No. 4 Penn State rallied from a 14-point deficit in the second half for a 33-30 victory over USC. Allar passed for a career-high 391 yards and two touchdowns despite three interceptions for Penn State (6-0, 3-0 Big Ten), which didn’t lead in the final 49 minutes of regulation. The Nittany Lions came back three times to tie it in the final 20 minutes, the last on Nicholas Singleton’s 14-yard TD reception with 2:53 left.

USC then got shoved 3 yards backward on its first three plays of overtime before Michael Lantz missed a 45-yard field goal attempt. Penn State advanced to the 19, and Barker’s teammates stormed the Coliseum field to celebrate after he buried the winner.

“I think the word ‘resilient’ was probably the best word to define our team today,” Penn State coach James Franklin said. “And it’s good to be able to go in and say, ‘Hey guys, we’re a second-half team.’ But I’d prefer not to say that anymore. I’d prefer to be a four-quarter team, a start-fast team, all of it.”

Quentin Joyner scored two early TDs for the Trojans (3-3, 1-3), who have lost three of four. USC hasn’t beaten a top-five team since the Trojans held off No. 5 Penn State in a 52-49 thriller of a Rose Bowl in January 2017.

“It’s a gut punch, no doubt about it,” said Lincoln Riley, who has lost eight of his last 13 games.

Penn State relied heavily on Warren, who racked up 224 yards and a touchdown even though his only catch after the third quarter was a 3-yarder in overtime. He matched the record for catches by a tight end set by New Mexico’s Emilio Vallez in 1967 and equaled by Jon Harvey for Northwestern in 1982.

“In the flow of the game, you don’t realize that,” Warren said, shrugging off his records. “I thought we just did an all-around good job.”